## Problem 184 — rank:281

**Title:** Persistence of intragenotypic variance

**Added binding assumptions (LaTeX):**

```latex
Maximizing a lineage growth exponent implies invasion resistance only when the competitor experiences the same exogenous environment and resident interactions do not change its growth law. Otherwise use an explicit resident-dependent invasion exponent. Specify the sampling law for phenotype variance, including ancestry, age and environmental history; conditioning only on current $e_t$ can mix persistent histories. Nonzero residual variance alone does not establish that variation is adaptive.
```

**Direct edits:** 0. **Fresh external checks:** None; supplied sources retained as provenance.