## Problem 310 — rank:292

**Title:** Extreme variation in genome size

**Added binding assumptions (LaTeX):**

```latex
For a fixed finite irreducible generator, solving the stationary linear equations is an established computation; on a countable space, positive recurrence and nonexplosion require proof. Specify how insertion/deletion jumps respect the lower size boundary and how lineage substitution rates relate to population-level fixation. A stationary distribution across evolutionary histories is not automatically the distribution of genome sizes among contemporaneous individuals or taxonomically selected species.
```

**Direct edits:** 0. **Fresh external checks:** None; supplied sources retained as provenance.